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  ― 117 ―

CCXLVIII. E. Lane-Poolei Maiden.

In Journ. Roy. Soc. N.S.W., liii, 107 (1919).

FOLLOWING is the original description:—

Arbor mediocris, White Gum vocata; cortice crassa, pulvere alba tecta; ligno hepatico; foliis primariis lanceolatis vel lato-lanceolatis, ca 6 cm. longis 3 cm. latis, venis secondariis fere parallellis; foliis maturis breve petioliatis, lanceolatis, acuminatis subfalcatis, ca 10 vel 11 cm. longis, 2 cm. latis, venis inconspicuis; pedunculis teretibus, ca 1·5 cm. longis, plerumque 4–6 floris, pedicellis, teretibus 1 cm. longis; calycis tubo ca. 1 cm. diametro, fere hemispherico; operculo crassissimo, hemispherico; antheris grandis fissuris parallelis late dehiscentibus; fructibus hemisphericis, ca. 1 cm. diametro, margine lato, leniter convexo, valvis distincte exsertis.

A medium-sized tree, known as “White Gum,” and carrying a thick bark covered with a white powder. Sapwood pale-coloured and thick, the timber interlocked, and rich reddish-brown in colour, drying, in the course of years, to a deep purplish-brown.

Juvenile leaves shortly petiolate, lanceolate to broadly-lanceolate, about 6 cm. long by 3 cm. broad, of the same colour on both sides, the secondary veins moderately spreading, and tending to be parallel to each other. A vein more prominent than the other secondary veins, roughly following the outline of the leaf, but at a considerable distance from the margin, and giving the leaf a triplinerved appearance.

Mature leaves shortly petiolate, lanceolate, acuminate, slightly falcate, not large, usually about 10 or 11 cm. long, and up to 2 cm. broad, venation inconspicuous, the fine veins roughly parallel and making an angle of about 45 degrees with the midrib, intramarginal vein hardly removed from the edge.

Peduncles axillary or lateral, terete, about 1·5 cm. long, bearing usually 4 to 6 moderately large flowers on terete pedicels up to 1 cm. long. Buds shiny.

Calyx-tube nearly hemispherical, about 1 cm. in diameter, with two slightly raised ridges separated by 180 degrees; tapering rather abruptly into the pedicel.

Operculum very thick, hemispherical or terminating in a slight but sharp point when nearly ripe. When less ripe, slightly broader than the calyx-tube, and without a point.

Stamens about 9 mm. long, inflected in the bud, anthers large, opening widely in parallel slits. Gland long, faintly visible at the back. Filament at the base. The anthers belong to the Platyantheræ group.

Disc broadish, oblique, forming a prominent ring round the ovary, of which the obtusely conical centre protrudes slightly above the disc at the time of flowering.

Fruit hemispherical, about 1 cm. in diameter, the rim broad, slightly convex, the capsule not sunk, the valves conspicuously exsert.

Type from Beenup, W.A. (C. E. Lane-Poole, No. 465).

Named in honour of Charles E. Lane-Poole, Conservator of Forests of Western Australia, who collected this species, and who has done much to promote the study of this genus in his State.




  ― 118 ―

Range.

It is confined to Western Australia, and, so far as is known at present, to a strip of coast-land, more or less ascending the Darling Range, in the south-western portion of the State, on the Perth-Bunbury Railway Line, between Kelmscott 16, and Waroona, 70 miles south of Perth.

“Very clean White Gum, Kelmscott, foot of Darling Range, 16 miles south of Perth.” (Dr. J. B. Cleland, No. 4.) Figured at fig. 4a and 4b, Plate 74, of the present work.

“White Gum,” 40 feet high, 12 inches in diameter, near Beenup, S.W. Railway, on the Perth to Bunbury road, 24 miles south of Perth (C. E. Lane-Poole, No. 8, November, 1918, fruits only; No. 465, July, 1919, complete material).

“A White Gum, sandy scrub land, Serpentine River, W.A.” In Herb. Melb., and variously attributed by Mueller (on the label) to E. uncinata and to E. micranthera.note It is a very old specimen, and is figured at fig. 8a, 8b, 8c of Plate 74 of the present work. This and the following three localities are practically identical.

“Salmon-white Gum or Powder Bark Wandoo. Height to about 40 feet, to 3 feet in diameter.” Near Keysbrook (39 miles south of Perth), near the Belgobin School, on the Perth-Bunbury road. (Mr. Schock, through C. E. Lane-Poole, under the same number, 8, as given to some Beenup specimens.)

Tree of 40 feet, 3 feet in diameter. Keysbrook, Perth-Bunbury road (Mr. Schock, per Dr. F. W. Stoward, No. 1).

“Salmon Gum or Powder-bark Wandoo,” half a mile south of Serpentine River on Perth-Bunbury road. (Mr. Schock, per Dr. F. W. Stoward, No. 90.)

Sent as “Wandoo,” Waroona, January, 1903 (Forester J. J. Fitzgerald). Waroona is 70 miles south of Perth, and I could only obtain buds. Referred to at p. 224, Part XVII of the present work.

AFFINITIES.

1. With E. redunca Schauer.

That officers of the Forest Department of Western Australia should, quite independently, in 1903 and 1918, speak of this as a Wandoo, shows that the general appearance of the tree, its bark and timber, must bear more than a superficial resemblance to the true Wandoo (E. redunca). But comparison of the figures 4 and 8, Plate 74 of this work, which partly depict E. Lane-Poolei, and Plate 140, which shows E. redunca, shows that the two species are botanically very dissimilar.




  ― 119 ―

2. With E. accedens W. V. Fitzgerald.

Mr. Schock, the Collector of the Forest Department of Western Australia, calls E. Lane-Poolei “Powder-bark Wandoo.” Both species are White Gums, with white, powdery barks, and the timbers have some external resemblance. The sylvicultural conditions of the two trees require to be worked out. As to the use of the term “Powder-bark,” Part XXXIV, p. 101, of this work may be referred to. If we turn to Plate 142 of the same Part, and compare it with figures 4 and 8 of Plate 74, which in part depict E. Lane-Poolei, it will be seen that the two species have no close botanical affinity.

3. With E. Oldfieldii F.v.M.

The affinity of this species is with E. Oldfieldii, which included E. Drummondii Benth., a species which in Part XVII of this work I erroneously followed Mueller in suppressing. I have shown, in Part XLI, how these two species differ. The affinity of E. Lane-Poolei is with E. Drummondii rather than with E. Oldfieldii sensu strictu. E. Oldfieldii is a Mallee with fruits in threes, while E. Lane-Poolei is a tree of considerable size, with fruits up to six in the head. The buds and leaves, both juvenile and mature, are very different.

4. With E. Drummondii Benth.

This species, of which but little is known in the field, is described from the York district as “a small tree of about 20 feet, with trunk and branches smooth, whitish-buff, with a few brown semi-detached scales of dead bark.” Additional field-notes are very desirable, but it would appear that E. Lane-Poolei is a different tree, and a Powder-bark.

Comparison, however, with figures 3, 5, 6, 7, 9, 10 of Plate 74 (E. Drummondii), together with a good specimen of the type of this species, is sufficient to show that it and E. Lane-Poolei (figures 4 and 8 of Plate 74) are sufficiently different. The leaves of E. Drummondii are commonly, perhaps preponderatingly, ovate-lanceolate, the buds more ovoid, with the opercula longer than the calyx-tube; the fruits are smaller, and very different.

5. With E. Campaspe S. le M. Moore.

It is interesting to note a resemblance in the very thick, hemispherical operculum of E. Campaspe, but the anthers are different, and so are the fruits and other characters. But one so frequently receives, particularly from distant places, botanical specimens which are quite fragmentary, and a hint which may put one on one's guard may be useful.

6. With E. oleosa F.v.M.

In its anthers it belongs to the Platyantheræ, which includes E. oleosa and its allies. The species are, however, very different in many other respects, but endeavour will be made to discuss these relationships when the seedlings of all the species are brought into comparison.

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